A comparative study is made of two groups of Chelicerata (Palpigradi and Actinotrichid mites, together constituting the Epimerata, a chelicerate class), and models of the evolution of epimerate characters are prepared. The study is based on the same methods and principles as the first part of the present series, and the same terminology has been used. The models of the evolution of characters constitute the epimerate archetype and the standard of epimerate classification.