A new species, Zygogynum vinkii, is described within the woody Ranalean family Winteraceae. It differs from other species of the endemic New Caledonian genus in having 302-371 stamens per flower. Unspecialized cells of the lower leaf surface are covered with cuticular papillae of a type hitherto unreported in Zygogynum. The three species with non-tetrad pollen, Z. pomiferum, Z. mackeei and Z. vinkii are compared.